[1] Guangdong Acad Sci, Inst Zool, Guangdong Key Lab Anim Conservat & Resource Utiliz, Guangdong Publ Lab Wild Anim Conservat & Utilizat, Guangzhou 510260, Peoples R China
[2] Smithsonian Inst, Dept Paleobiol, Natl Museum Nat Hist, Washington, DC 20013 USA
[3] Capital Normal Univ, Coll Life Sci & Acad Multidisciplinary Studies, Beijing 100048, Peoples R China
[4] Univ Maryland, Dept Entomol, College Pk, MD 20742 USA
[5] Guangzhou Coll Technol & Business, Sch Engn, Guangzhou 510850, Peoples R China