[1] Chinese Acad Sci, Xinjiang Inst Ecol & Geog, Xinjiang Key Lab Desert Plant Roots Ecol & Vegetat, Urumqi 830011, Peoples R China
[2] Chinese Acad Sci, Xinjiang Inst Ecol & Geog, State Key Lab Desert & Oasis Ecol, Urumqi 830011, Peoples R China
[3] Cele Natl Stn Observat & Res Desert Grassland Ecos, Cele 848300, Peoples R China
[4] Univ Chinese Acad Sci, Beijing 100049, Peoples R China
[5] Guizhou Univ, Coll Agr, Coll Life Sci, Guiyang 550025, Peoples R China
[6] King Khalid Univ, Coll Sci, Biol Dept, Abha 61421, Saudi Arabia
[7] King Khalid Univ, Appl Coll, POB 9004, Abha 61413, Saudi Arabia
[8] King Khalid Univ, Ctr Bee Res & Its Prod CBRP, POB 9004, Abha 61413, Saudi Arabia