[1] Chinese Acad Sci, Northwest Inst Ecoenvironm & Resources, Key Lab Ecol Safety & Sustainable Dev Arid Lands, Urat Desert Grassland Res Stn, Lanzhou, Peoples R China
[2] Key Lab Stress Physiol & Ecol, Lanzhou 730000, Gansu Province, Peoples R China
[3] Chinese Acad Sci, Xinjiang Inst Ecol & Geog, State Key Lab Desert & Oasis Ecol, Urumqi 830011, Peoples R China
[4] North Wyke, Rothamsted Res, Devon EX20 2SB, England