[1] Lund Univ, Dept Geol Quaternary Sci, Lund, Sweden
[2] Cty Museum Norrbotten, Lulea, Sweden
[3] Cty Museum Jonkoping, Jonkoping, Sweden
[4] Lanzhou Univ, Coll Earth & Environm Sci, Key Lab Western Chinas Environm Sci, Minist Educ, Lanzhou, Peoples R China
[5] Chinese Acad Sci, Inst Tibetan Plateau Res, State Key Lab Tibetan Plateau Earth Syst Resources, Beijing, Peoples R China
[6] Univ Chinese Acad Sci, Coll Resources & Environm, Beijing, Peoples R China
[7] Univ Cambridge, Dept Zool, Cambridge, England
[8] Univ Copenhagen, Globe Inst, Ctr Ancient Environm Genom, Copenhagen, Denmark