[1] Nelson Mandela Univ, Inst Coastal & Marine Res, POB 77000, ZA-6031 Gqeberha, South Africa
[2] CSIR, Stellenbosch Pretoria, South Africa
[3] South African Natl Biodivers Inst, Kirstenbosch Res Ctr, Private Bag X7, ZA-7735 Claremont, South Africa
[4] Univ Cape Town, Dept Biol Sci, ZA-7701 Rondebosch, South Africa
[5] Univ Witwatersrand, Restorat & Conservat Biol Res Grp, Sch Anim Plant & Environm Sci, Private Bag 3, ZA-2050 Johannesburg, South Africa
[6] Univ Pretoria, Dept Geog Geoinformat & Meteorol Geog, ZA-0083 Pretoria, South Africa
[7] Dept Agr Forestry & Fisheries, Fisheries Res & Dev, Cape Town, South Africa
[8] Nelson Mandela Univ, Inst Coastal & Marine Res, DSI NRF Res Chair Shallow Water Ecosyst, Dept Bot, POB 77000, ZA-6031 Gqeberha, South Africa
[9] SAAMBR, Oceanog Res Inst, Durban, South Africa
[10] Univ KwaZulu Natal, Sch Life Sci, Durban, South Africa