[1] Univ Bristol, Fac Life Sci, Sch Cellular & Mol Med, Colorectal Tumour Biol Grp, Biomed Sci Bldg,Univ Walk, Bristol BS8 1TD, England
[2] Canc Res UK Beatson Inst, Garscube Estate, Switchback Rd, Bearsden G61 1BD, Glasgow, Scotland
[3] Cardiff Univ, Sch Med, Div Canc & Genet, Cardiff CF14 4XN, Wales
[4] Univ Sussex, Sch Life Sci, Sussex House, Brighton BN1 9RH, England
[5] Monash Univ Biomed Discovery Inst, Melbourne 3800, Australia