[1] Murdoch Univ, Coll Sci Hlth Engn & Educ, Sch Vet Med, Murdoch, WA 6150, Australia
[2] Univ Utrecht, GD Anim Hlth, Deventer, NL-3584 CS Utrecht, Netherlands
[3] Univ Utrecht, Fac Vet Med, NL-3584 CS Utrecht, Netherlands
[4] Anexa Vet Serv, Cognosco, Morrinsville 3340, New Zealand
[5] Univ Melbourne, Fac Vet & Agr Sci, Anim Welf Sci Ctr, Parkville, Vic 3052, Australia
[6] Univ Melbourne, Fac Vet & Agr Sci, Werribee, Vic 3030, Australia
[7] Murdoch Univ, Inst Immunol & Infect Dis, Murdoch, WA 6150, Australia
[8] Murdoch Univ, Future Foods Inst, Ctr Anim Prod & Hlth, Murdoch, WA 6150, Australia
[9] Murdoch Univ, Harry Butler Inst, Murdoch, WA 6150, Australia