[1] Jianshui Research Station, School of Soil and Water Conservation, Beijing Forestry University, Beijing,100083, China
[2] State Key Laboratory of Vegetation and Environmental Change, Zhejiang Qianjiangyuan Forest Biodiversity National Observation and Research Station, Institute of Botany, Chinese Academy of Sciences, Beijing,100093, China
[3] Key Laboratory of Biodiversity Conservation of National Forestry and Grassland Administration, Ecology and Nature Conservation Institute, Chinese Academy of Forestry, Beijing,100091, China