[1] Key Laboratory of Desert and Desertification, Cold and Arid Regions Environmental and Engineering Research Institute, Chinese Academy of Sciences, Lanzhou-730000, Gansu, China
[2] Key Laboratory of Extreme Environmental Microbial Resources and Engineering, Lanzhou-730000, Gansu, China
[3] State Key Laboratory of Cryospheric Sciences, Cold and Arid Regions Environmental and Engineering Research Institute, Chinese Academy of Sciences, Lanzhou-730000, Gansu, China
[4] University of Chinese Academy of Sciences, Beijing,100049, China