[1] College of Life Sciences, Zhejiang Normal University, Jinhua,321004, China
[2] Guangdong Key Lab of Geodynamics and Geohazards, School of Earth Sciences and Engineering, Sun Yat-sen University, Zhuhai,519082, China
[3] Southern Marine Science and Engineering Guangdong Laboratory (Zhuhai), Zhuhai,519082, China
[4] Guangxi Laboratory on the Study of Coral Reef in the South China Sea, Coral Reef Research Centre of China, School of Marine Sciences, Guangxi University, Nanning,530004, China
[5] Center for Ocean-Atmosphere Interaction Research, Nanjing University of Information Science and Technology, Nanjing,210044, China
[6] State Key Laboratory of Estuarine and Coastal Research, East China Normal University, Shanghai,200241, China
[7] Institute of Evolutionary Sciences of Montpellier, University of Montpellier, Montpellier,34000, France
[8] School of Geography, South China Normal University, Guangzhou,510635, China