*Telephone: +86-25-84399690. Fax: +86-25-84395314. E-mail: zhanglin2012@njau.edu.cn. ORCID Lin Zhang: 0000-0003-1555-1086 Funding This work was supported by the National Natural Science Foundation of China (31402094 and 31601957);
the Fundamental Research Funds for the Central Universities of China (KYZ201641);
the National Key Research and Development Program of China (2016YFD0500501);
and the Three Agricultural Projects of Jiangsu Province (SXGC2017281). Notes The authors declare no competing financial interest;