[1] Department of Neuroscience, Physiology and Pharmacology, Division of Biosciences, University College London, London,WC1E 6BT, United Kingdom
[2] Department of Synthetic Biology and Immunology, National Institute of Chemistry, Hajdrihova 19, Ljubljana,1000, Slovenia
[3] Vollum Institute, Oregon Health & Science University, Portland,OR,97239, United States
[4] Howard Hughes Medical Institute, Oregon Health & Science University, Portland,OR,97239, United States