[1] Department of Bioscience and Bioinformatics, Kyushu Institute of Technology, Fukuoka, Iizuka,820–8502, Japan
[2] PRESTO, Japan Science and Technology Agency, Saitama, Kawaguchi,332–0012, Japan
[3] Department of Molecular Cell Biology, Faculty of Science, VU University Amsterdam, Amsterdam,1081 HZ, Netherlands