[1] Natl Inst Oceanog & Fisheries, NIOF, Cairo, Egypt
[2] Univ Hassan II Casablanca, Fac Sci Ain Chock, Dept Earth Sci, Km 8 Route El Jadida BP 5366, Casablanca 20100, Morocco
[3] Natl Agcy Land Conservat Cadastre & Cartog, Rabat, Morocco
[4] Tanta Univ, Fac Sci, Bot & Microbiol Dept, Tanta 31512, Egypt
[5] Smithsonian Environm Res Ctr, Edgewater, MD 21037 USA