[1] Wageningen Univ & Res, Dept Environm Technol, Wageningen, Netherlands
[2] Wageningen Univ & Res, Lab Microbiol, Wageningen, Netherlands
[3] Delft Univ Technol, Dept Geosci & Engn, Delft, Netherlands
[4] IE Univ, Dept Environm Sci Sustainabil, Segovia, Spain