[1] Cadi Ayyad Univ, Fac Sci Semlalia, Dept Biol, Lab Agrofood Biotechnol & Valorizat Plant Bioresou, Marrakech 40000, Morocco
[2] Hassan II Univ Casablanca, Fac Sci & Techn Mohammedia, Dept Biol, Lab Biochem Environm & Agrifood URAC 36, Mohammadia 28800, Morocco
[3] Mohammed Ist Univ, Multidisciplinary Fac Nador, Dept Biol, Nador 62700, Morocco
[4] Fac Sci & Tech, Dept Biol, Lab Environm & Hlth, Errachidia 52000, Morocco
[5] Cadi Ayyad Univ, Ctr Agrobiotechnol & Bioingn, CNRST Ctr Agrobiotech URL CNRST 05, Unite Rech Labellisee, Marrakech 40000, Morocco