[1] Nanchang Univ, Sch Life Sci, Nanchang 330031, Jiangxi, Peoples R China
[2] Univ Melbourne, Fac Sci, Dept Vet Biosci, Parasitol Res Grp, Parkville, Vic 3010, Australia
[3] South China Agr Univ, Coll Life Sci, Coll Agr, Guangdong Lab Lingnan Modern Agr,State Key Lab Con, Guangzhou 510642, Peoples R China
[4] Zhejiang Museum Nat Hist, Hangzhou 310012, Zhejiang, Peoples R China