[1] Univ Lincoln, Coll Hlth & Sci, Sch Life & Environm Sci, Dept Life Sci, Brayford Pool Campus, Lincoln LN6 7TS, England
[2] Univ Lincoln, Coll Hlth & Sci, Sch Psychol, Lincoln, England
[3] CNR, Ist Sci & Tecnol Cogniz, Rome, Italy
[4] Univ Abertay Dundee, Div Psychol & Forens Sci, Dundee, Scotland
[5] Univ Portsmouth, Ctr Comparat & Evolutionary Psychol, Dept Psychol, Portsmouth PO1 2DY, England