[1] Russian Acad Sci, Ctr Forest Ecol & Prod, Moscow 117997, Russia
[2] Russian Acad Sci, Inst Systemat & Ecol Anim, Siberian Branch, Novosibirsk 630091, Russia
[3] Siberian Fed Univ, Dept Ecol & Nat Management, Krasnoyarsk 660041, Russia
[4] Russian Acad Sci, VN Sukachev Inst Forest, Siberian Branch, Krasnoyarsk 660036, Russia
[5] Russian Acad Sci, Inst Biophys, Siberian Branch, Krasnoyarsk 660036, Russia