[1] Univ Seville, Agron Dept, ETSIA, Crta Utrera Km 1, Seville 41013, Spain
[2] Unit associated CSIC Uso Sostenible Suelo & Agua A, Crta Utrera Km 1, E-41013 Seville, Spain
[3] Univ Politecn Madrid, Agr Prod Dept, CEIGRAM, Ave Puerta Hierro 2, Madrid 28040, Spain
[4] Inst Recursos Nat & Agrobiol Sevilla IRNAS CSIC, Ave Reina Mercedes 10, Seville 41012, Spain