[1] Chulalongkorn Univ, Fac Sci, Dept Biol, Integrat Insect Ecol Res Unit, Bangkok, Thailand
[2] South African Museum, Res & Exhibit Dept, Iziko Museums South Afr, POB 61, ZA-8000 Cape Town, South Africa
[3] Univ Cape Town, Dept Biol Sci, Private Bag, ZA-7701 Rondebosch, South Africa
[4] Tel Aviv Univ, Steinhardt Museum Nat Hist, IL-69978 Tel Aviv, Israel
[5] Uppsala Univ, Museum Evolut, Zool, Norbyvagen 16, SE-75236 Uppsala, Sweden