[1] Inner Mongolia Univ, Sch Ecol & Environm, Minist Educ, Key Lab Ecol & Resource Use Mongolian Plateau, Hohhot 010021, Peoples R China
[2] Chinese Acad Sci, Inst Appl Ecol, Key Lab Forest Ecol & Management, Shenyang 110016, Peoples R China
[3] Univ Chinese Acad Sci, Coll Resources & Environm, Beijing 100049, Peoples R China
[4] Inner Mongolia Autonomous Reg Ecol & Agr Meteorol, Hohhot 010051, Peoples R China
[5] Minist Educ China & Inner Mongolia Autonomous Reg, Collaborat Innovat Ctr Grassland Ecol Secur, Hohhot 010021, Peoples R China
[6] Aarhus Univ, Ctr Ecol Dynam Novel Biosphere ECONOVO, Dept Biol, Ny Munkegade 114, DK-8000 Aarhus C, Denmark
[7] Aarhus Univ, Ctr Biodivers Dynam Changing World BIOCHANGE, Dept Biosci, Ny Munkegade 114, DK-8000 Aarhus C, Denmark