[1] US Forest Serv, USDA, Madison, WI 53726 USA
[2] Univ Amsterdam, Inst Biodivers & Ecosyst Dynam, Amsterdam, Netherlands
[3] Princeton Univ, High Meadows Environm Inst, Princeton, NJ USA
[4] Univ Hawaii Manoa, Hawaii Inst Marine Biol, Honolulu, HI USA
[5] Oregon State Univ, Coll Earth Ocean & Atmospher Sci, Corvallis, OR USA