[1] Hokkaido Univ, Int Inst Zoonosis Control, Div Risk Anal & Management, Sapporo, Japan
[2] Hokkaido Univ, Fac Vet Med, Dept Dis Control, Lab Parasitol, Sapporo, Japan
[3] Hokkaido Univ, Grad Sch Vet Med, Dept Clin Sci, Lab Vet Internal Med, Sapporo, Japan
[4] Hokkaido Univ, Fac Vet Med, Dept Dis Control, Lab Mol Med, Sapporo, Japan
[5] Hokkaido Inst Publ Hlth, Dept Infect Dis, Sapporo, Hokkaido, Japan