[1] Univ Fed Goias, Dept Ecol, Programa Posgrad Ecol & Evolucao, Campus Samambaia, BR-74690900 Goiania, GO, Brazil
[2] Univ Fed Goias, Inst Ciencias Biol, Dept Ecol, Campus Samambaia, BR-74690900 Goiania, GO, Brazil
[3] Univ Fed Campinas, Dept Biol Anim, BR-13083970 Campinas, SP, Brazil
[4] Univ Fed Paraiba, Dept Biociencias, BR-58397000 Areia, PB, Brazil