[1] De La Salle Univ, Coll Sci, Dept Biol, Manila, Philippines
[2] De La Salle Univ, Ctr Nat Resources & Environm Res, Plant & Soil Hlth Res Unit, Manila, Philippines
[3] Univ Idaho, Parma Res & Extens Ctr, Parma, ID 83660 USA
[4] USDA ARS, Small Grains & Potato Germplasm Res Unit, Aberdeen, ID USA