[1] Inst Nacl Pesquisas Amazonia INPA, Lab Sistemat & Ecol Coleoptera LASEC, Ave Bem-Te-Vi,8-406-Campus II,Petropolis, BR-69067001 Manaus, AM, Brazil
[2] Zool Inst, Russian Acad Sci, Lab Insect Systemat, Univ nab 1, St Petersburg 199034, Russia
[3] Western Carolina Univ, Biol Dept, 535 Apodaca Bldg,1 Univ Dr, Cullowhee, NC 28723 USA