[1] Museum Koenig, Leibniz Inst Anal Biodivers Change, D-53113 Bonn, Germany
[2] Helmholtz Ctr Environm Res GmbH, Dept Conservat Biol, UFZ, D-04318 Leipzig, Germany
[3] Comenius Univ, Dept Zool, Bratislava, Slovakia
[4] Univ Rostock, Inst Biosci, Gen & Systemat Zool, D-18055 Rostock, Germany