[1] Guangxi Univ, Coral Reef Res Ctr China, Sch Marine Sci, Guangxi Lab Study Coral Reefs South China Sea, Nanning 530004, Peoples R China
[2] Nanjing Hongchuang Explorat Technol Serv, Nanjing 210023, Peoples R China
[3] China Geol Survey, Nanjing Ctr, Nanjing 210016, Peoples R China
[4] Guangxi Acad Sci, Nanning 530007, Peoples R China
[5] Inst Beibu Gulf Marine Ind, Fangchenggang 538000, Peoples R China