[1] Chinese Acad Sci, CAS, Qingdao, Peoples R China
[2] Chinese Acad Sci, Inst Oceanol, Ctr Ocean Mega Sci, Shandong Prov Key Lab Expt Marine Biol, Qingdao, Peoples R China
[3] Qingdao Natl Lab Marine Sci & Technol, Lab Marine Biol & Biotechnol, Qingdao, Peoples R China
[4] Univ Chinese Acad Sci, Beijing 100049, Peoples R China
[5] Qingdao Agr Univ, Coll Life Sci, Qingdao 266109, Peoples R China